Abstract
The Aegean Sea is highly impacted by biological invasions with 209 alien species being reported until the end of 2019, of which 149 are considered as established. Among these, the most abundant group is Pisces, followed by Mollusca, Crustacea, Annelida, and Macrophyta. There is an increasing trend of new species introductions in the Aegean Sea, reaching 60 new introductions between 2010 and 2019. The most common pathway of introduction in the Aegean Sea is “unaided” introduction from neighboring ecoregions, i.e., natural dispersal across the Aegean borders, without human agency. Most of these species are Lessepsian immigrants, introduced through the Suez Canal, and initially recorded in the Levantine Sea. The second most common pathway in the Aegean is “transport-stowaway,” i.e., introduction through ballast waters, hull fouling, or with angling/fishing equipment. All other pathways are of much lower importance for the Aegean Sea. The most invasive species in the region are the fish Siganus rivulatus, Siganus luridus, Fistularia commersonii, Lagocephalus sceleratus, and Pterois miles; the macroalgae Caulerpa cylindracea, Codium fragile, Stypopodium schimperi, Asparagopsis taxiformis, and Womersleyella setacea; the crab Callinectes sapidus; and potentially the molluscs Conomurex persicus, Pinctada imbricata radiata, Dendostrea cf. folium, and Fulvia fragilis.
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Acknowledgments
Stelios Katsanevakis was supported by the Hellenic Foundation for Research and Innovation (H.F.R.I.) under the “First Call for H.F.R.I. Research Projects to support Faculty members and Researchers and the procurement of high-cost research equipment grant” (Project ALAS – “ALiens in the Aegean – a Sea under siege” [108]; Project Number: HFRI-FM17-1597).
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Annex 1
Annex 1
Alien species in the Aegean Sea, sorted by the year of first record in the Aegean. Their taxonomic, establishment success, and pathway of introduction in the Aegean are also provided. Pathways: UNA unaided, TS transport-stowaway, TC transport-contaminant, COR corridor, REL release in nature, EC escape from confinement, UNK unknown
Taxonomic group | Species | Establishment success | Pathway | Year of first record |
---|---|---|---|---|
Macroalgae/Rhodophyta | Hypnea cornuta (Kützing) J. Agardh | Established | TS | 1894 |
Chordata/Ascidiacea | Ascidiella aspersa (Müller, 1776) | Established | UNK | 1901 |
Chordata/Ascidiacea | Ciona robusta Hoshino and Tokioka, 1967 (as Ciona intestinalis (Linnaeus, 1767)) | Established | TS | 1901 |
Macroalgae/Rhodophyta | Lophocladia lallemandii (Montagne) Schmitz | Established | TS | 1908 |
Tracheophyta | Halophila stipulacea (Forsskål) Ascherson | Established | CORR | 1923 |
Chordata/Pisces | Siganus rivulatus Forsskål, 1775 | Established | UNA | 1925 |
Macroalgae/Ochrophyta | Cutleria multifida (Turner) Greville | Established | TS | 1932 |
Chordata/Pisces | Sargocentron rubrum (Forsskål, 1775) | Established | UNA | 1940 |
Cnidaria/Scyphozoa | Cassiopea andromeda (Forsskål, 1775) | Established | UNA | 1942 |
Chordata/Pisces | Hemiramphus far (Forsskål, 1775) | Established | UNA | 1943 |
Chordata/Pisces | Stephanolepis diaspros Fraser-Brunner, 1940 | Established | UNA | 1943 |
Chordata/Pisces | Equulites klunzingeri (Steindachner, 1898) | Casual | UNA | 1946 |
Chordata/Pisces | Parexocoetus mento (Valenciennes, 1846) | Established | UNA | 1946 |
Crustacea/Decapoda | Callinectes sapidus Rathbun, 1896 | Established | TS | 1947 |
Chordata/Pisces | Upeneus moluccensis (Bleeker, 1855) | Established | UNA | 1947 |
Chordata/Pisces | Lagocephalus guentheri Miranda Ribeiro, 1915 | Established | UNA | 1952 |
Foraminifera | Coscinospira hemprichii Ehrenberg, 1839 | Established | UNA, TS | 1955 |
Annelida | Ficopomatus enigmaticus (Fauvel, 1923) | Established | TS | 1959 |
Chordata/Pisces | Alepes djedaba (Forsskål, 1775) | Established | UNA | 1960 |
Mollusca/Bivalvia | Pinctada imbricata radiata (Leach, 1814) | Established | REL | 1961 |
Crustacea/Stomatopoda | Erugosquilla massavensis (Kossmann, 1880) | Established | UNA | 1963 |
Annelida | Notomastus aberans Day, 1957 | Established | UNA | 1964 |
Annelida | Paradyte cf. crinoidicola (Potts, 1910) | Casual | UNA | 1964 |
Chordata/Pisces | Siganus luridus (Rüppell, 1829) | Established | UNA | 1964 |
Foraminifera | Amphistegina lobifera (Larsen, 1976) | Established | UNA | 1967 |
Crustacea/Copepoda | Arietellus pavoninus (G. O. Sars, 1905) | Established | UNA, TS | 1967 |
Bryozoa | Crisularia serrata (Lamarck, 1816) | Unknown | TS | 1967 |
Bryozoa | Microporella coronata (Audouin, 1826) | Questionable | TS | 1967 |
Macroalgae/Ochrophyta | Pylaiella littoralis (Linnaeus) Kjellman | Questionable | TS | 1967 |
Chordata/Ascidiacea | Styela plicata (Lesueur, 1823) | Established | TS | 1968 |
Bryozoa | Amathia verticillata (delle Chiaje, 1822) | Established | TS | 1969 |
Mollusca/Gastropoda | Nerita sanguinolenta Menke, 1829 | Casual | UNA | 1969 |
Crustacea/Cirripedia | Amphibalanus eburneus (Gould, 1841) | Unknown | TS | 1970 |
Crustacea/Cirripedia | Balanus trigonus Darwin, 1854 | Established | TS | 1970 |
Chordata/Pisces | Saurida lessepsianus Russell, Golani, and Tikochinski, 2015 | Established | UNA | 1971 |
Mollusca/Bivalvia | Brachidontes pharaonis (P. Fischer, 1870) | Established | UNA, TS | 1975 |
Mollusca/Gastropoda | Bursatella leachii (De Blainville, 1817) | Established | UNA, TS | 1975 |
Annelida | Lysidice collaris Grube, 1870 | Established | UNA | 1975 |
Annelida | Protodorvillea biarticulata Day, 1963 | Established | TS | 1975 |
Annelida | Sigambra parva (Day, 1963) | Established | TS | 1975 |
Annelida | Axionice medusa (Savigny in Lamarck, 1818) | Established | UNA | 1976 |
Annelida | Dodecaceria capensis Day, 1961 | Established | TS | 1976 |
Annelida | Hydroides elegans (Haswell, 1883) | Established | TS | 1976 |
Macroalgae/Rhodophyta | Hypnea spinella (C. Agardh) Kützing | Established | TS | 1979 |
Macroalgae/Rhodophyta | Ceramium bisporum Ballantine | Casual | TS | 1980 |
Macroalgae/Rhodophyta | Chondria collinsiana Howe | Questionable | TS | 1980 |
Macroalgae/Rhodophyta | Sarconema scinaioides Børgesen | Casual | EC | 1980 |
Cnidaria/hydrozoa | Clytia linearis (Thornely, 1900) | Established | UNA | 1981 |
Macroalgae/Ochrophyta | Padina boryana Thivy | Questionable | UNA | 1981 |
Chordata/Pisces | Apogonichthyoides pharaonis (Bellotti, 1874) | Established | UNA | 1982 |
Mollusca/Gastropoda | Conomurex persicus (Swainson, 1821) | Established | UNA, TS | 1983 |
Chordata/Pisces | Pempheris rhomboidea Kossmann and Rauber, 1877 | Established | UNA | 1983 |
Annelida | Polycirrus twisti Potts, 1928 | Established | UNA | 1983 |
Crustacea/Decapoda | Thalamita poissonii (Audouin, 1826) | Established | UNA | 1983 |
Mollusca/Bivalvia | Mya arenaria Linnaeus, 1758 | Established | TS, TC | 1984 |
Mollusca/Gastropoda | Crepidula fornicata (Linnaeus, 1758) | Established | TC, TS | 1985 |
Mollusca/Bivalvia | Petricolaria pholadiformis Lamarck, 1818 | Established | TS | 1985 |
Chordata/Pisces | Atherinomorus forskalii (Rüppell, 1838) | Established | UNA | 1986 |
Macroalgae/Ochrophyta | Colpomenia peregrina Sauvageau | Established | TS | 1986 |
Bryozoa | Crepidacantha poissoni (Audouin, 1826) | Unknown | TS | 1986 |
Annelida | Desdemona ornata Banse, 1957 | Casual | TS | 1986 |
Mollusca/Gastropoda | Rapana venosa (Valenciennes, 1846) | Casual | UNA | 1986 |
Foraminifera | Heterostegina depressa d’Orbigny, 1826 | Established | UNA | 1988 |
Crustacea/Copepoda | Paracartia grani (G. O. Sars, 1904) | Established | TS | 1988 |
Macroalgae/Rhodophyta | Womersleyella setacea (Hollenberg) Norris | Established | TS | 1988 |
Annelida | Branchiomma luctuosum (Grube, 1869) | Established | TS | 1989 |
Annelida | Chaetozone corona Berkeley and Berkeley, 1941 | Established | TS | 1989 |
Ctenophora | Mnemiopsis leidyi (Agassiz, 1865) | Established | UNA | 1990 |
Cnidaria/hydrozoa | Sertularia marginata (Kirchenpauer, 1864) | Established | TS | 1990 |
Mollusca/Gastropoda | Acteocina mucronata (Philippi, 1849) | Casual | UNA | 1991 |
Macroalgae/Chlorophyta | Caulerpa racemosa var. lamourouxii f. requienii (Montagne) Weber van Bose | Established | UNA | 1991 |
Annelida | Prionospio pulchra Imajima, 1990 | Casual | TC | 1991 |
Macroalgae/Rhodophyta | Asparagopsis taxiformis (Delile) Trevisan de Saint-Léon | Established | UNA | 1992 |
Macroalgae/Chlorophyta | Codium fragile (Suringar) Hariot | Established | TS | 1992 |
Chordata/Pisces | Pteragogus trispilus Randall, 2013 | Established | UNA | 1992 |
Mollusca/Bivalvia | Anadara transversa (Say, 1822) | Established | TS | 1993 |
Echinodermata | Ophiactis savignyi (Müller and Troschel, 1842) | Casual | UNA | 1993 |
Mollusca/Gastropoda | Smaragdia souverbiana (Montrouzier, 1863) | Established | UNA | 1993 |
Foraminifera | Spiroloculina antillarum d’Orbigny, 1839 | Established | UNA | 1993 |
Mollusca/Gastropoda | Cerithiopsis tenthrenois (Melvill, 1896) | Casual | UNA | 1994 |
Mollusca/Gastropoda | Melibe viridis (Kelaart, 1858) | Established | UNA, TS | 1994 |
Crustacea/Decapoda | Penaeus pulchricaudatus Stebbing, 1914 | Established | UNA | 1995 |
Chordata/Pisces | Planiliza haematocheila (Temminck and Schlegel, 1845) | Established | UNA | 1995 |
Chordata/Pisces | Sphyraena chrysotaenia Klunzinger, 1884 | Established | UNA | 1995 |
Macroalgae/Chlorophyta | Caulerpa cylindracea Sonder | Established | UNA | 1996 |
Chordata/Ascidiacea | Diplosoma listerianum (Milne Edwards, 1841) | Established | TS | 1996 |
Bryozoa | Hippopodina sp. A as Hippopodina feegeensis (Busk, 1884) | Established | TS | 1996 |
Mollusca/Bivalvia | Malleus regula (Forsskål in Niebuhr, 1775) | Established | UNA | 1996 |
Mollusca/Gastropoda | Mnestia girardi (Audouin, 1826) | Established | UNA | 1996 |
Macroalgae/Ochrophyta | Stypopodium schimperi (Buchinger ex Kützing) Verlaque and Boudouresque | Established | UNA | 1996 |
Mollusca/Bivalvia | Fulvia fragilis (Forsskål in Niebuhr, 1775) | Established | TS | 1997 |
Annelida | Spirorbis marioni Caullery and Mesnil, 1897 | Established | TS | 1997 |
Crustacea | Paradella dianae (Menzies, 1962) | Established | TS | 1997 |
Crustacea/Decapoda | Alpheus rapacida (de Man, 1908) | Casual | UNA | 1998 |
Mollusca/Gastropoda | Bulla arabica Malaquias and Reid, 2008 | Established | TS | 1998 |
Crustacea/Decapoda | Charybdis (Goniohellenus) longicollis Leene, 1938 | Established | UNA | 1999 |
Crustacea/Decapoda | Ixa monodi Holthuis and Gottlieb, 1956 | Established | UNA | 1999 |
Crustacea/Decapoda | Metapenaeopsis aegyptia Galil and Golani, 1990 | Established | UNA | 1999 |
Annelida | Leiocapitellides analis Hartmann-Schröder, 1960 | Casual | UNA | 2000 |
Crustacea/Decapoda | Portunus segnis (Forsskål, 1775) | Established | UNA | 2000 |
Chordata/Pisces | Fistularia commersonii (Rüppell, 1835) | Established | UNA | 2001 |
Foraminifera | Planogypsina acervalis (Brady, 1884) | Established | UNK | 2001 |
Chordata/Pisces | Callionymus filamentosus Valenciennes, 1837 | Established | UNA | 2003 |
Chordata/Pisces | Etrumeus golani Di Battista, Randall and Bowen, 2012 | Established | UNA | 2003 |
Chordata/Pisces | Lagocephalus suezensis Clark and Gohar, 1953 | Established | UNA | 2003 |
Annelida | Pseudonereis anomala (Gravier, 1900) | Established | TS | 2003 |
Chordata/Pisces | Sphyraena flavicauda Rüppell, 1838 | Established | UNA | 2003 |
Annelida | Spirobranchus tetraceros (Schmarda, 1861) | Established | TS | 2003 |
Chordata/Pisces | Tylosurus crocodilus Péron and Lesueur, 1821 | Casual | UNA | 2003 |
Chordata/Pisces | Upeneus pori Ben-Tuvia and Golani, 1989 | Established | UNA | 2003 |
Ctenophora | Beroe ovata Mayer, 1912 | Unknown | UNA | 2004 |
Mollusca/Gastropoda | Goniobranchus annulatus (Eliot, 1904) | Established | UNA, TS | 2004 |
Crustacea/Decapoda | Myra subgranulata Galil and Golani, 1990 | Established | UNA | 2004 |
Crustacea/Decapoda | Percnon gibbesi (H. Milne Edwards, 1853) | Established | UNA | 2004 |
Chordata/Pisces | Petroscirtes ancylodon Rüppell, 1838 | Established | UNA | 2004 |
Chordata/Pisces | Tylerius spinosissimus (Regan, 1908) | Established | UNA | 2004 |
Crustacea/Decapoda | Calappa pelii Herklots, 1851 | Casual | UNK | 2005 |
Crustacea/Decapoda | Coleusia signata (Paulson, 1875) | Established | UNA | 2005 |
Mollusca/Gastropoda | Haminoea cyanomarginata Heller and Thompson, 1983 | Established | UNA, TS | 2005 |
Chordata/Pisces | Lagocephalus sceleratus (Gmelin, 1788) | Established | UNA | 2005 |
Annelida | Pseudopolydora paucibranchiata (Okuda, 1937) | Established | TS | 2005 |
Echinodermata | Synaptula reciprocans (Forsskål, 1775) | Established | UNA | 2005 |
Macroalgae/Rhodophyta | Botryocladia madagascariensis G. Feldmann | Established | TS | 2006 |
Annelida | Caulleriella viridis (Langerhans, 1881) | Questionable | TS | 2006 |
Macroalgae/Rhodophyta | Melanothamnus harveyi (Bailey) Díaz-Tapia and Maggs | Established | TS | 2006 |
Foraminifera | Miliolinella fichteliana d’Orbigny, 1839 | Established | UNA | 2006 |
Annelida | Timarete punctata (Grube, 1859) | Casual | TS | 2006 |
Chordata/Pisces | Torquigener flavimaculosus Hardy and Randall, 1983 | Established | UNA | 2006 |
Mollusca/Bivalvia | Chama asperella Lamarck, 1819 | Established | TS | 2007 |
Mollusca/Gastropoda | Ergalatax junionae Houart, 2008 | Established | UNA | 2007 |
Annelida | Glycinde bonhourei Gravier, 1904 | Casual | UNA | 2007 |
Annelida | Linopherus canariensis Langerhans, 1881 | Casual | TC | 2007 |
Annelida | Neanthes agulhana (Day, 1963) | Casual | TS | 2007 |
Mollusca/Bivalvia | Crassostrea/Magallana sp./spp. | Established | EC/REL | 2008 |
Macroalgae/Rhodophyta | Hypnea anastomosans Papenfuss, Lipkin and Silva | Established | UNA | 2008 |
Annelida | Lepidonotus tenuisetosus (Gravier, 1902) | Casual | UNA | 2008 |
Chordata/Ascidiacea | Phallusia nigra Savigny, 1816 | Established | UNA | 2008 |
Annelida | Polydora cornuta Bosc, 1802 | Established | TS | 2008 |
Chordata/Pisces | Scomberomorus commerson Lacepède, 1800 | Established | UNA | 2008 |
Mollusca/Bivalvia | Spondylus cf. spinosus Schreibers, 1793 | Casual | TS | 2008 |
Chordata/Pisces | Terapon theraps Cuvier, 1829 | Casual | UNK | 2008 |
Mollusca/Bivalvia | Chama pacifica Broderip, 1834 | Established | UNA | 2009 |
Mollusca/Gastropoda | Coryphellina rubrolineata O’Donoghue, 1929 | Established | UNA | 2009 |
Macroalgae/Rhodophyta | Laurencia caduciramulosa Masuda and Kawaguchi | Established | TS | 2009 |
Crustacea/Decapoda | Macrophthalmus indicus Davie, 2012 | Casual | UNA | 2009 |
Crustacea/isopoda | Paracerceis sculpta (Holmes, 1904) | Established | TS | 2009 |
Crustacea/Copepoda | Parvocalanus crassirostris (Dahl, 1894) | Established | UNK | 2009 |
Chordata/Pisces | Pterois miles (Bennett, 1828) | Established | UNA | 2009 |
Mollusca/Cephalopoda | Sepioteuthis lessoniana Férussac in Lesson, 1831 complex | Established | UNA | 2009 |
Macroalgae/Rhodophyta | Asparagopsis armata Harvey | Casual | UNA | 2010 |
Crustacea/Decapoda | Charybdis (Charybdis) hellerii (A. Milne-Edwards, 1867) | Established | UNA | 2010 |
Mollusca/Bivalvia | Dendostrea cf. folium (Linnaeus, 1758) | Established | UNA, TS | 2010 |
Chordata/Pisces | Lutjanus sebae (Cuvier, 1816) | Casual | EC | 2010 |
Crustacea/Cirripedia | Megabalanus tintinnabulum (Linnaeus, 1758) | Casual | TS | 2010 |
Mollusca/Bivalvia | Septifer cumingii (Dunker, 1855) | Established | UNA | 2010 |
Crustacea/Decapoda | Atergatis roseus (Rüppell, 1830) | Established | UNA | 2011 |
Crustacea/Decapoda | Carupa tenuipes Dana, 1851 | Established | UNA | 2011 |
Crustacea/Copepoda | Centropages furcatus (Dana, 1852) | Established | UNA | 2011 |
Mollusca/Gastropoda | Cerithium scabridum Philippi, 1848 | Established | UNA | 2011 |
Crustacea/Decapoda | Gonioinfradens giardi (Nobili, 1905) | Established | UNA | 2011 |
Crustacea/Amphipoda | Caprella scaura Templeton, 1836 | Established | UNK | 2012 |
Chordata/Pisces | Champsodon nudivittis (Ogilby, 1895) | Established | UNA | 2012 |
Foraminifera | Euthymonacha polita (Chapman, 1900) | Casual | UNK | 2012 |
Crustacea/isopoda | Paranthura japonica Richardson, 1909 | Established | TS | 2012 |
Crustacea/Decapoda | Penaeus aztecus Ives, 1891 | Established | UNA | 2012 |
Crustacea/Decapoda | Penaeus hathor (Burkenroad, 1959) | Established | UNA | 2012 |
Mollusca/Gastropoda | Syrnola fasciata Jickeli, 1882 | Established | UNA | 2012 |
Macroalgae/Chlorophyta | Caulerpa taxifolia var. distichophylla (Sonder) Verlaque, Huisman and Procaccini | Established | UNA | 2013 |
Mollusca/Gastropoda | Diodora funiculata (Reeve, 1850) | Established | TS | 2013 |
Annelida | Leonnates persicus Wesenberg-Lund, 1949 | Casual | UNA | 2013 |
Mollusca/Gastropoda | Pyrunculus fourierii (Audouin, 1826) | Casual | UNA | 2013 |
Macroalgae/Ochrophyta | Scytosiphon dotyi Wynne | Questionable | TS | 2013 |
Crustacea/Decapoda | Xanthias lamarckii (H. Milne Edwards, 1834) | Casual | UNA | 2013 |
Chordata/Pisces | Bregmaceros nectabanus Whitley, 1941 | Established | UNA | 2014 |
Porifera | Paraleucilla magna Klautau, Monteiro and Borojevic, 2004 | Established | TC | 2014 |
Mollusca/Gastropoda | Syphonota geographica (A. Adams and Reeve, 1850) | Established | UNA | 2014 |
Crustacea/Amphipoda | Bemlos leptocheirus (Walker, 1909) | Unknown | UNA | 2015 |
Annelida | Branchiomma bairdi (McIntosh, 1885) | Established | TS | 2015 |
Bryozoa | Celleporaria brunnea (Hincks, 1884) | Unknown | TS | 2015 |
Bryozoa | Celleporaria vermiformis (Waters, 1909) | Established | TS | 2015 |
Mollusca/Gastropoda | Cerithiopsis pulvis (A. Issel, 1869) | Casual | UNA | 2015 |
Crustacea/isopoda | Cymodoce fuscina Schotte and Kensley, 2005 | Casual | TS | 2015 |
Crustacea/Decapoda | Dyspanopeus sayi (Smith, 1969) | Casual | TS | 2015 |
Chordata/Ascidiacea | Herdmania momus (Savigny, 1816) | Established | UNA | 2015 |
Annelida | Hydroides brachyacantha Rioja, 1941 | Casual | TS | 2015 |
Annelida | Hydroides dirampha Mörch, 1863 | Established | TS | 2015 |
Mollusca/Gastropoda | Rhinoclavis kochi (Philippi, 1848) | Established | UNA | 2015 |
Mollusca/Gastropoda | Sticteulima sp. [cf. lentiginosa (A. Adams, 1861)] | Casual | UNA, TS | 2015 |
Chordata/Ascidiacea | Symplegma brakenhielmi (Michaelsen, 1904) | Established | TS | 2015 |
Bryozoa | Tricellaria inopinata d’Hondt and Occhipinti Ambrogi, 1985 | Unknown | TS | 2015 |
Echinodermata | Diadema setosum (Leske, 1778) | Established | UNA | 2016 |
Mollusca/Bivalvia | Isognomon legumen (Gmelin, 1791) | Established | UNA | 2016 |
Crustacea/isopoda | Mesanthura cf. romulea Poore and Lew-Ton, 1986 | Unknown | TS | 2016 |
Mollusca/Gastropoda | Oscilla galilae Bogi, Karhan and Yokeş, 2012 | Casual | UNA | 2016 |
Chordata/Pisces | Synchiropus sechellensis Regan, 1908 | Established | UNA | 2016 |
Mollusca/Gastropoda | Viriola sp.[cf. bayani] Jousseaume, 1884 | Established | UNA | 2016 |
Chordata/Pisces | Abudefduf sexfasciatus (Lacepède, 1801) | Casual | EC | 2017 |
Chordata/Pisces | Acanthurus sohal (Forsskål, 1775) | Casual | CORR | 2017 |
Chordata/Pisces | Scarus ghobban Forsskål in Niebuhr, 1775 | Established | UNA | 2017 |
Mammalia | Sousa plumbea (G. Cuvier, 1829) | Casual | UNA | 2017 |
Crustacea/isopoda | Sphaeroma walkeri Stebbing 1905 | Unknown | TS | 2017 |
Cnidaria/Scyphozoa | Chrysaora cf. achlyos Martin, Gershwin, Burnett, Cargo and Bloom, 1997 | Casual | TS | 2018 |
Tracheophyta | Halophila decipiens Ostenfeld | Casual | TS | 2018 |
Crustacea/Copepoda | Oithona davisae Ferrari F.D. and Orsi, 1984 | Casual | UNA/TS | 2018 |
Chordata/Pisces | Parupeneus forsskali (Fourmanoir and Guézé, 1976) | Established | UNA | 2018 |
Chordata/Pisces | Lutjanus argentimaculatus (Forsskål, 1775) | Casual | UNA | 2019 |
Chordata/Pisces | Pomadasys stridens (Forsskål, 1775) | Unknown | UNA | 2019 |
Cnidaria/Scyphozoa | Rhopilema nomadica Galil, 1990 | Established | UNA | 2019 |
Chordata/Pisces | Vanderhorstia mertensi Klausewitz, 1974 | Casual | UNA | 2019 |
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Katsanevakis, S., Zenetos, A., Corsini-Foka, M., Tsiamis, K. (2020). Biological Invasions in the Aegean Sea: Temporal Trends, Pathways, and Impacts. In: The Handbook of Environmental Chemistry. Springer, Berlin, Heidelberg. https://doi.org/10.1007/698_2020_642
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DOI: https://doi.org/10.1007/698_2020_642
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